Plantago major L.

First published in Sp. Pl.: 112 (1753)
This species is accepted
The native range of this species is Temp. Eurasia to Arabian Peninsula, Macaronesia, N. & S. Africa. It is an annual or perennial and grows primarily in the temperate biome. It is has environmental uses, as animal food and a medicine and for food.

Descriptions

Plantaginaceae, B. Verdcourt. Flora of Tropical East Africa. 1971

Morphology General Habit
An extremely variable, usually robust, glabrous or pubescent herb, with short stout erect stem and numerous adventitious roots.
Morphology Leaves
Leaves borne in a rosette, spirally arranged; blade ovate, elliptic or rarely rounded, (1-)10- 20(-30) cm. long. (0.7-)5-9.5(-17) cm. wide, rounded at the apex, entire, sinuate or irregularly toothed, 3-9-nerved at the base, abruptly narrowed at the base into a petiole usually ± equalling the blade, 1-20 cm. long.
Morphology Reproductive morphology Inflorescences
Inflorescences spicate, the spikes nearly always cylindric, almost invariably simple, (0.5)10-15(-70) cm. long; peduncles (l-)7-15(-40) cm. long; bracts 1-2.5 mm. long, acute, brownish with a brown keel.
Morphology Reproductive morphology Flowers Calyx
Sepals broadly elliptic to rounded, 1.5-2 mm. long and wide, keeled.
Morphology Reproductive morphology Flowers Corolla
Corolla greenish- or yellowish-white, 2-4 mm. long; lobes elliptic-ovate, ovate or narrowly triangular, 1-1.25 mm. long, obtuse or acute.
Morphology Reproductive morphology Flowers Androecium Stamens Anthers
Anthers at first lilac, later whitish or yellowish.
Morphology Reproductive morphology Fruits
Capsule globose or subconic, 2-3 mm. long, 3-34-seeded.
Morphology Reproductive morphology Seeds
Seeds brown to dark brown or olivaceous with lighter belt on back, ellipsoidal or angular, 1-1.7 mm. long, 0.8 mm. wide, the ventral side more convex than the dorsal side; hilum lighter.
Figures
Fig. 1/1, 2.
Habitat
Grassy river-banks; 1665 m.
Distribution
K4 ranging naturally throughout Europe and northern and central Asia, but now naturalized throughout most of the world
[FTEA]

Bernal, R., G. Galeano, A. Rodríguez, H. Sarmiento y M. Gutiérrez. 2017. Nombres Comunes de las Plantas de Colombia. http://www.biovirtual.unal.edu.co/nombrescomunes/

Vernacular
chiracru, chiracu, llantén
[UNAL]

The Useful Plants of Boyacá project

Ecology
Alt. 100 - 2600 m.
Morphology General Habit
Herb.
Conservation
Least concern.
Distribution
Naturalised in Colombia.
[UPB]

IUCN Red List of Threatened Species https://www.iucnredlist.org/species/168960/1257878

Conservation
LC - least concern
[IUCN]

Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá. http://catalogoplantasdecolombia.unal.edu.co

Distribution
Naturalizada en Colombia; Alt. 100 - 2600 m.; Amazonia, Andes, Sierra Nevada de Santa Marta.
Morphology General Habit
Hierba
Conservation
Preocupación Menor
[CPLC]

Plantaginaceae, G. Lehmann. Flora Zambesiaca 9:1. 1988

Morphology General Habit
Perennial herb, very variable, usually robust, glabrous or pubescent, growing with a short straight erect stem, with numerous adventitious roots.
Morphology Leaves
Leaves in a basal rosette spirally arranged, petiolate; lamina (1.5) 5–30 (40) × (0.5) 3–10 (17) cm., ovate to elliptic or rarely subcircular, entire sinuate or irregularly dentate, 3–9-veined at the base, usually glabrous or rarely pubescent, with the apex rounded to subobtuse, abruptly narrowed into the petiole; petiole 3–38 cm. long, usually as long as the lamina or shorter, cauliculate, flexible, glabrous or rarely pubescent.
Morphology Reproductive morphology Inflorescences Scape
Scape (1) 7–15 (47) cm. long, 1–4 mm. in diam., erect or ascending, striate, glabrous or sometimes with short, appressed or ascending hairs.
Morphology Reproductive morphology Inflorescences
Inflorescence spicate, the spikes nearly always cylindrical, almost invariably simple, (0.5) 10–15 (70) cm. long, 3–8 mm. in diam.; flowers densely to loosely arranged.
Morphology Reproductive morphology Inflorescences Bracts
Bracts 1–2 (2.5) mm. long, ovate, glabrous, midrib distinct.
Morphology Reproductive morphology Flowers Calyx
Sepals 1.5–2 (2.5) mm. long, almost boat-shaped, usually equal, glabrous, green with narrow scarious margins, midrib prominent.
Morphology Reproductive morphology Flowers Corolla
Corolla tube c. 2 mm. long, glabrous; lobes 1–1.25 mm. long, lanceolate to ovate or narrowly triangular, acute to subacute, glabrous.
Morphology Reproductive morphology Flowers Androecium Stamens
Stamens 2–3 mm. long, exserted; anthers at first lilac, later whitish or yellowish.
Morphology Reproductive morphology Flowers Gynoecium Pistil
Ovary globose; pistil longer than the flower, hairy.
Morphology Reproductive morphology Fruits
Capsule 2–3 (5) mm. long, ovoid to globose or subconical, (3) 8–16 (34) seeded.
Morphology Reproductive morphology Seeds
Seeds 1–1.7 mm. long, ellipsoid or ellipsoid-trigonous, with the ventral side more convex than the dorsal.
[FZ]

Ghazanfar, S. A. & Edmondson, J. R (Eds). (2014) Flora of Iraq, Volume 5 Part 2: Lythraceae to Campanulaceae.

Morphology General Habit
Perennial herb, acaulous, (10–)20–40(–70) cm tall with a short rootstock and many fibrous roots
Morphology Leaves
Leaves large, elliptic-ovate to orbicular-ovate, (5–)10–25(–40) × (1–) 4–10 (–20) cm, base tapering into petiole about as long as lamina, apex rounded, margins entire or irregularly dentate or sinuate-dentate, strongly 5–9-veined, glabrous or rough with sparse minute hairs
Morphology Reproductive morphology Inflorescences Scape
Scape erect, ascending or arcuate, equal to leaves or shorter, pilose with sparse minute soft hairs
Morphology Reproductive morphology Inflorescences
Spike dense or lax, linear or linear-cylindrical, (8–)5–30(–50) cm; bract ovate, as long as or usually shorter than calyx, 1–2 mm, along midrib herbaceous and slightly keeled, scarious at margins, entire, obtuse, glabrous
Morphology Reproductive morphology Flowers Calyx
Sepals ovate-elliptic to orbicular, 1.5– 2.5 mm, with a distinct midrib, margins scarious, obtuse, glabrous
Morphology Reproductive morphology Flowers Corolla
Corolla greenish or yellowish, lobes ovate, 0.5–1 mm long, obtuse or acute
Morphology Reproductive morphology Fruits
Capsule longer than calyx, globose or conical, tapering above, 2-celled with 4–30 seeds
Morphology Reproductive morphology Seeds
Seeds angular, smooth.
Ecology
In wet shaded places in farms and orchards, muddy places amongst rocks, near banks of shallow ponds and streams, in rich humus soil and on rich soil with limestone in upland forests; alt.: 150–900m
Phenology
Flowering and fruiting: Mar.–Aug.
Distribution
Throughout Iraq except in the desert and plateau region; found in the upper plains and foothills and the alluvial plains. Europe to Siberia, from the Mediterranean region, Sinai, Arabian Peninsula to Iran and India; naturalized in N America and many other parts of the world.
Vernacular
BIZIR DINBIL (recorded by Blakelock, loc.cit.); TARAQUZ (recorded on label by Guest).
[FIQ]

Morphology Reproductive morphology Seeds Protein Content

19.10% Entire seed/nut. Moisture content not stated (Barclay & Earle, 1974)

[SID]

Distribution
Biogeografic region: Amazonia, Andean. Elevation range: 100–2600 m a.s.l. Naturalised in Colombia. Colombian departments: Amazonas, Antioquia, Bogotá DC, Boyacá, Cundinamarca, Huila, Magdalena, Meta, Nariño, Putumayo, Valle del Cauca.
Habit
Herb.
Conservation
IUCN Red List Assessment (2021): LC.
Ecology
Habitat according IUCN Habitats Classification: forest and woodland, savanna, shrubland, desert, artificial - terrestrial.
[UPFC]

Extinction risk predictions for the world's flowering plants to support their conservation (2024). Bachman, S.P., Brown, M.J.M., Leão, T.C.C., Lughadha, E.N., Walker, B.E. https://nph.onlinelibrary.wiley.com/doi/full/10.1111/nph.19592

Conservation
Predicted extinction risk: not threatened. Confidence: confident
[AERP]

Uses

Use Animal Food
Seeds - The seeds are eaten by birds (Florez-Cárdenas et al. 2010).
Use Medicines Digestive System Disorders
Leaves - Used in liquid medicines (Florez-Cárdenas et al. 2010).
Use Medicines Injuries
Leaves - Used in poultices (Florez-Cárdenas et al. 2010). Leaves - Used in poultices to promote healing (Lagos-López 2007, Florez-Cárdenas et al. 2010).
Use Medicines Pain
Leaves - Used as an analgesic (Florez-Cárdenas et al. 2010).
Use Medicines Skin or Subcutaneous Cellular Tissue Disorders
Leaves - Used in the treatment of acne (Florez-Cárdenas et al. 2010).
Use Medicines Unspecified Medicinal Disorders
Medicinal (State of the World's Plants 2016, Instituto Humboldt 2014).
[UPB]

Use
A species of some economic value whose seeds and leaves are used medicinally as a laxative and a coolant; also used for application on boils. Several subspecific ranks were recognized in the original manuscript based mainly on leaf characters. I found it difficult to assign every collection to the subspecific rank recognized as many intermediates exist. The species is best treated as a single polymorphic species. The var., subvar. and f. are listed in synonymy.
[FIQ]

Use Animal Food
Used as animal food.
Use Environmental
Environmental uses.
Use Gene Sources
Used as gene sources.
Use Food
Used for food.
Use Materials
Used as material.
Use Medicines
Medical uses.
[UPFC]

Common Names

English
Plantain, Red-leaved Plantain
Spanish
Llantén, plantén, plantain.

Sources

  • Angiosperm Extinction Risk Predictions v1

    • Angiosperm Threat Predictions
    • http://creativecommons.org/licenses/by/4.0
  • Catálogo de Plantas y Líquenes de Colombia

    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Flora Zambesiaca

    • Flora Zambesiaca
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Flora of Iraq

    • Flora of Somalia
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Flora of Tropical East Africa

    • Flora of Tropical East Africa
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Herbarium Catalogue Specimens

    • Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/
  • IUCN Categories

    • IUCN Red List of Threatened Species
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Kew Living Collection Database

    • Common Names from Plants and People Africa http://www.plantsandpeopleafrica.com/
  • Kew Names and Taxonomic Backbone

    • The International Plant Names Index and World Checklist of Vascular Plants 2024. Published on the Internet at http://www.ipni.org and https://powo.science.kew.org/
    • © Copyright 2023 International Plant Names Index and World Checklist of Vascular Plants. http://creativecommons.org/licenses/by/3.0
  • Kew Science Photographs

    • Copyright applied to individual images
  • Seed Information Database

    • Royal Botanic Gardens Kew. (2019) Seed Information Database (SID). Version 7.1. Available from: http://data.kew.org/sid/ (September 2019)
    • http://creativecommons.org/licenses/by/3.0
  • Universidad Nacional de Colombia

    • ColPlantA database
    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Useful Plants and Fungi of Colombia

    • http://creativecommons.org/licenses/by-nc-sa/3.0
  • Useful Plants of Boyacá Project

    • ColPlantA database
    • http://creativecommons.org/licenses/by-nc-sa/3.0
    • http://creativecommons.org/licenses/by/3.0/
  • World Checklist of Vascular plants (WCVP)

    • The International Plant Names Index and World Checklist of Vascular Plants 2024. Published on the Internet at http://www.ipni.org and https://powo.science.kew.org/
    • © Copyright 2023 World Checklist of Vascular Plants. http://creativecommons.org/licenses/by/3.0